@charset "UTF-8";@import url("https://fonts.googleapis.com/css2?family=Raleway:ital,wght@0,300;0,400;0,600;0,700;1,500&family=Source+Sans+Pro:ital,wght@0,300;0,400;0,700;1,300;1,400;1,700&display=swap");@layer tokens, base, layout, components, utilities;@layer tokens{:root{--color-primary:#368093;--color-link:#53a3b7;--color-text:#576674;--color-heading:#687683;--color-heading-hero:#8a949d;--color-muted:#6E7881;--color-muted-text:#6b6b6b;--color-breadcrumb-link:#4a5a68;--color-heading-link-hover:#0290b3;--color-small:#999;--gray-50:#F6F7F8;--gray-100:#eceff1;--gray-200:#C7CFD6;--gray-300:#a3b0bb;--gray-400:#687683;--gray-500:#4c5966;--gray-600:#576674;--neutral-100:#f7f7f7;--neutral-150:#f5f5f5;--neutral-200:#f2f2f2;--neutral-250:#ececec;--neutral-300:#e7e7e7;--neutral-350:#e4e4e4;--neutral-400:#d8d8d8;--neutral-500:#ccc;--neutral-600:#777;--neutral-900:#252525;--danger:#b94a48;--danger-bg:#f2dede;--danger-border:#eed3d7;--danger-strong:#a33;--warning-text:#7a5e00;--info-bg:#F0F8FF;--highlight-bg:#FFFFD4;--well-bg:#eaeef2;--distmap-border:#cfd8dc;--distmap-pre2000:#90a4ae;--distmap-post2000:#1976d2;--distmap-empty-text:#546e7a;--distmap-empty-icon:#b0bec5;--distmap-error:#b71c1c;--footer-text:#dbe2e9;--footer-meta:#c1cfdb;--header-bg:#4c5966;--nav-text:#d8dfe5;--nav-hover:#53a3b7;--nav-sub-bg:#758694;--nav-sub-rule:#a3b0bb;--nav-panel-bg:#6d767e;--footer-link:#9bd9e9;--footer-credits-bg:#576674;--footer-credits-rule:#a3b0bb;--hr-dot:#c8d2da;--rule-light:#d8dfe5;--focus-ring:#368093;--search-trigger-text:#758694;--font-body:"Source Sans Pro",sans-serif;--font-heading:"Raleway",sans-serif;--container-max:1100px;--gutter:2rem;--radius:3px;--radius-sm:2px;--space-1:0.4rem;--space-2:0.8rem;--space-3:1.2rem;--space-4:1.6rem;--space-5:2.4rem;--space-6:3.2rem}}@layer base{html{margin:0;padding:0;font-size:62.5%;-webkit-font-smoothing:antialiased;-webkit-text-size-adjust:100%;text-size-adjust:100%}body{margin:0;padding:0;overflow-x:hidden;background:#fff;color:var(--color-text);font-family:var(--font-body);font-size:15px;line-height:1.6}@media (min-width:980px){body{line-height:1.65}}@media (max-width:600px){body{font-size:16px;line-height:1.5}}article,aside,details,figcaption,figure,footer,header,main,nav,section{display:block}h1,h2,h3,h4,h6{margin:2% 0;font-family:var(--font-heading);font-weight:normal;color:var(--color-heading);text-rendering:optimizeLegibility}h1 a,h2 a,h3 a,h4 a,h6 a{color:inherit}h1 a:focus-visible,h1 a:hover,h2 a:focus-visible,h2 a:hover,h3 a:focus-visible,h3 a:hover,h4 a:focus-visible,h4 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img{display:block;margin-inline:auto}figcaption{clear:both;margin-top:0;padding:0;background:var(--gray-50);font-size:90%;font-style:italic;text-align:left}table{max-width:100%;border-collapse:collapse;border-spacing:0;background-color:transparent}button,input,label,select,textarea{margin:0;font-family:var(--font-body);font-size:15px;font-weight:normal;line-height:20px;vertical-align:middle}label{display:block;margin-bottom:5px}button,input[type=button],input[type=checkbox],input[type=radio],input[type=reset],input[type=submit],label,select{cursor:pointer}button,input[type=button],input[type=reset],input[type=submit]{appearance:button}input[type=date],input[type=email],input[type=number],input[type=password],input[type=search],input[type=text],input[type=url],select,textarea{display:inline-block;box-sizing:content-box;height:20px;padding:4px 6px;margin-bottom:10px;line-height:20px;color:var(--color-text);vertical-align:middle;background-color:#fff;border:1px solid var(--gray-200);border-radius:4px;box-shadow:inset 0 1px 1px rgba(0,0,0,0.075);transition:border-color 0.2s linear,box-shadow 0.2s linear}input[type=date]:focus,input[type=email]:focus,input[type=number]:focus,input[type=password]:focus,input[type=search]:focus,input[type=text]:focus,input[type=url]:focus,select:focus,textarea:focus{outline:0;border-color:var(--color-link);box-shadow:inset 0 1px 1px rgba(0,0,0,0.075),0 0 0 3px rgba(83,163,183,0.25)}input,textarea{width:206px}input[type=date],input[type=email],input[type=number],input[type=password],input[type=search],input[type=text],input[type=url]{font-size:14px}input[type=search]{appearance:textfield}textarea{height:auto;overflow:auto;vertical-align:top;resize:vertical}input[type=checkbox],input[type=radio]{width:auto;margin:4px 0 0;line-height:normal}input[type=radio]{vertical-align:text-bottom}input[type=button],input[type=file],input[type=image],input[type=reset],input[type=submit]{width:auto}@media (prefers-reduced-motion:reduce){*{animation-duration:0.01ms!important;animation-iteration-count:1!important;transition-duration:0.01ms!important;scroll-behavior:auto!important}}@media print{*{background:transparent!important;color:#000!important;box-shadow:none!important;text-shadow:none!important}a,a:visited{text-decoration:underline}blockquote,pre{border:1px solid var(--neutral-600);page-break-inside:avoid}blockquote,blockquote p{padding-left:0!important}blockquote:before{display:none}thead{display:table-header-group}img,tr{page-break-inside:avoid}img{max-width:100%!important}h2,h3,p{orphans:3;widows:3}h2,h3{page-break-after:avoid}h1,h2,h3,h4,h6{margin:10px 0!important}ul{margin:0}hr{height:0;border-top:1px solid var(--neutral-500);background:none}}}@layer layout{#main-content{clear:both;padding-block:0 2%}.wrap{width:min(90%,var(--container-max));margin-inline:auto}.grid{display:grid;grid-template-columns:repeat(12,minmax(0,1fr));gap:var(--gutter)}.grid>*{grid-column:span 12;min-width:0}@media (min-width:768px){.col-4{grid-column:span 4}.col-7{grid-column:span 7}.col-8{grid-column:span 8}}.cluster{display:flex;flex-wrap:wrap;gap:var(--cluster-gap,var(--space-2) var(--space-3));align-items:var(--cluster-align,center)}.with-sidebar{display:flex;flex-wrap:wrap;gap:var(--gutter)}.with-sidebar>:first-child{flex-basis:0;flex-grow:999;min-inline-size:60%}.with-sidebar>:last-child{flex-basis:var(--sidebar-width,30rem);flex-grow:1}.card-grid{display:grid;grid-template-columns:repeat(auto-fill,minmax(var(--card-min,16rem),1fr));gap:var(--card-gap,var(--gutter));margin:0;padding:0;list-style:none}}@layer components{.page-hero{position:relative;clear:both;margin:-1% 0 0;padding:2% 0;font-size:1.8rem;font-weight:200;line-height:3rem;text-shadow:0 1px 1px rgba(255,255,255,0.7)}.page-hero__title{margin:0;padding-bottom:2rem;font-family:var(--font-heading);font-size:200%;font-weight:300;line-height:1.2;color:var(--gray-300);background-image:linear-gradient(to right,var(--hr-dot) 1px,transparent 1px);background-size:3px 1px;background-repeat:repeat-x;background-position:0 100%}.page-hero--center{text-align:center}.page-hero--no-border .page-hero__title{background-image:none;color:var(--color-heading-hero)}.page-hero--tight .page-hero__title{padding-bottom:0}.page-hero--midpage{margin-top:1em}@media (min-width:980px){.page-hero__title{font-size:250%}}@media (min-width:1100px){.page-hero__title{font-size:280%}}@media (max-width:767px){.page-hero{padding:2% 0}.page-hero__title{font-size:160%;line-height:1.3}}@media (max-width:500px){.page-hero{padding:2% 0 3%}.page-hero__title{font-weight:500}}@media (min-width:768px){.page-hero__title--small{font-size:220%}}@media print{.page-hero{margin:0;padding:0}.page-hero__title{background-image:none}}.section-title{margin:0 0 1em;padding-bottom:0.9rem;border-bottom:1px solid var(--rule-light);font-family:var(--font-heading);font-weight:500;line-height:normal;text-transform:lowercase}.section-title>span{padding-bottom:1rem;border-bottom:5px solid var(--rule-light)}.section-title--propercase{text-transform:none}*+.section-title{margin-top:3%}.button{display:inline-flex;align-items:center;gap:var(--space-1);padding:var(--space-1) var(--space-3);border:1px solid var(--color-primary);border-radius:var(--radius);background:var(--color-primary);color:#fff;font:inherit;line-height:1.5;text-decoration:none;cursor:pointer}.button:focus-visible,.button:hover{background:var(--color-link);border-color:var(--color-link);color:#fff;text-decoration:none}.button--subtle{background:var(--color-muted);border-color:var(--color-muted)}.button--ghost{background:var(--neutral-200);border-color:var(--neutral-400);color:var(--color-text)}.button--ghost:focus-visible,.button--ghost:hover{background:var(--color-muted);border-color:var(--color-muted);color:#fff}.button--small{padding:0.2rem var(--space-2);font-size:85%}.button svg{flex:none}.search-combo{display:flex;max-width:100%}.search-combo>input{flex:1 1 auto;min-width:0;margin:0;padding:var(--space-1) var(--space-2);border:1px solid var(--neutral-500);border-right:0;border-radius:var(--radius) 0 0 var(--radius);font:inherit;font-size:16px;line-height:1.5;color:var(--color-text);background:#fff}.search-combo>.button,.search-combo>button{flex:none;border-radius:0 var(--radius) var(--radius) 0}.search-combo>.button,.search-combo>button,.search-combo>input{height:3.6rem;padding-block:0;box-sizing:border-box}.card{display:flex;flex-direction:column;gap:var(--space-2);min-width:0}.card>a{display:block}.card img{display:block;width:100%;height:auto}.card__title{margin:0;font-family:var(--font-heading);font-weight:500;font-size:100%;line-height:1.3}.disclosure{border-top:1px solid var(--rule-light);border-bottom:1px solid var(--rule-light)}.disclosure>summary{padding:var(--space-2) 0;cursor:pointer;font-family:var(--font-heading);font-weight:500;list-style:none}.disclosure>summary::-webkit-details-marker{display:none}.disclosure>summary:after{content:"";display:inline-block;width:0.6em;height:0.6em;margin-left:var(--space-2);border-right:2px solid currentColor;border-bottom:2px solid 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var(--rule-light);font-family:var(--font-heading);font-size:1.8rem;font-weight:500;line-height:normal;text-transform:lowercase}.widget-box__title>span{padding-bottom:1rem;border-bottom:5px solid var(--rule-light)}.widget-box p{font-size:90%}.widget-box>form:last-child,.widget-box>p:last-child{margin-bottom:0}.widget-box ul{margin:0 0 var(--space-3);padding:0;list-style:none}.widget-box ul li{position:relative;padding:0.3rem 0 0.3rem 1.3rem}.widget-box ul li:before{content:"";position:absolute;left:0;top:1.1rem;width:0.5rem;height:0.5rem;border-right:1px solid currentColor;border-top:1px solid currentColor;transform:rotate(45deg);color:var(--gray-300)}.widget-box ul a{display:block;text-decoration:none}.widget-box ul a:hover,.widget-box ul li.active>a{color:var(--color-link)}.widget-box ul li.active:before,.widget-box ul li:hover:before{color:var(--color-link)}.widget-box ul ul{margin:0 0 0.5rem 1rem;font-size:90%}@media (max-width:767px){.widget-box{padding:var(--space-4);background:var(--gray-50);margin-bottom:var(--space-3)}}.card--row{flex-direction:row;flex-wrap:wrap;align-items:flex-start;gap:var(--space-3) var(--space-4);margin-bottom:1em;padding:var(--space-3);background:var(--gray-50)}.card--row .card__media{flex:0 0 auto;max-width:28%}.card--row .card__media img{width:auto;max-width:100%}.card--row .card__body{flex:1 1 24rem;min-width:0;display:flex;flex-direction:column;gap:var(--space-2)}.card--row .card__body>*{margin:0}.card--row .card__title{font-size:2rem;font-weight:300;color:var(--color-link)}.card--row .card__title a{text-decoration:none}.card--row .card__title strong{font-weight:600}.card--row .card__actions{align-self:flex-end;margin-top:auto}.comment{margin-bottom:3rem}.comment__heading{margin:0 0 1em;padding-bottom:0.9rem;border-bottom:1px solid var(--rule-light);font-family:var(--font-heading);font-size:1.6rem;font-weight:500;line-height:normal;text-transform:lowercase}.comment__heading>span{padding-bottom:1rem;border-bottom:5px solid var(--rule-light)}.comment__heading>small,.comment__heading>time{font-size:85%;font-weight:400;color:var(--color-muted)}.comment__heading>.comment__flag{float:right;font-weight:600;text-transform:none}.comment__body{padding:2%;background:var(--well-bg)}.comment__body>:last-child{margin-bottom:0}.comment__reply{margin-top:1rem;padding:0.5rem 1rem;border:1px solid var(--neutral-350);background:var(--neutral-100);font-size:90%;font-style:italic}.comment__reply p{margin-bottom:0.5rem}.comment__edit{display:block;text-align:right;font-size:85%}.pager-nav{margin:var(--space-5) 0;text-align:center}.pager-nav__list{display:inline-flex;flex-wrap:wrap;justify-content:center;margin:0;padding:0;list-style:none;border:1px solid var(--neutral-400);border-radius:var(--radius);overflow:hidden}.pager-nav li{margin:0;border-left:1px solid var(--neutral-400)}.pager-nav li:first-child{border-left:0}.pager-nav a,.pager-nav>ul>li>span{display:block;padding:var(--space-1) var(--space-3);line-height:2.4rem;text-decoration:none}.pager-nav a:focus-visible,.pager-nav a:hover{background:var(--gray-50)}.pager-nav .is-current a{background:var(--color-primary);color:#fff;cursor:default}.pager-nav .MarkupPagerNavSeparator{padding:var(--space-1) var(--space-2);line-height:2.4rem;color:var(--color-muted)}.widget-search{margin:0}.widget-search .search-combo>input{font-size:1.4rem}.card-grid--tiles{grid-template-columns:repeat(2,minmax(0,1fr));gap:var(--space-3);margin-bottom:3rem}@media (min-width:600px){.card-grid--tiles{grid-template-columns:repeat(3,minmax(0,1fr))}}@media (min-width:900px){.card-grid--tiles{grid-template-columns:repeat(4,minmax(0,1fr))}}.card--tile{gap:0;background:var(--neutral-200);text-align:center;overflow:hidden}.card--tile .card__media{display:block}.card--tile .card__title{padding:3%;border-top:1px solid var(--neutral-300);font-family:var(--font-heading);font-size:1.2rem;font-weight:500;line-height:1.4}.card--tile .card__title em{font-style:italic}.card--tile a{text-decoration:none}.card--tile .card__media:focus-visible img,.card--tile:hover .card__media img{opacity:0.85}.table-scroll{overflow-x:auto;-webkit-overflow-scrolling:touch;margin-bottom:3rem}.data-table{width:100%;min-width:56rem;margin:0;border-collapse:collapse;border:1px solid var(--neutral-400);font-size:1.4rem}.data-table td,.data-table th{padding:var(--space-2);border-left:1px solid var(--neutral-400);border-top:1px solid var(--neutral-400);text-align:left;vertical-align:top}.data-table th{font-weight:600}.data-table tbody tr:nth-child(odd){background:var(--gray-50)}.data-table .data-table__id{width:clamp(60px,80px,10vw)}.data-table .data-table__bf{width:clamp(50px,65px,14vw)}.data-table .data-table__photo{width:clamp(100px,340px,42vw)}.search-form{max-width:48rem;margin:0 0 var(--space-4)}.search-form__label{display:block;margin-bottom:var(--space-1)}.search-results__count{margin:var(--space-4) 0 var(--space-3)}.highlight,mark{background:var(--highlight-bg);color:inherit;padding:0 0.1em}.month-link{margin-bottom:2rem}.species-list__items{margin:0 0 var(--space-4);padding:0;list-style:none}.species-list__item{margin:0;padding:0.5rem;font-size:1.6rem}.species-list__item:nth-child(odd){background:rgba(163,176,187,0.1)}@media (max-width:600px){.species-list__item{line-height:1.2;margin-bottom:1.5em}}.species-list__id{display:inline-block;min-width:10.8rem}.species-list__family-title{margin:0 0 1em;padding-bottom:0.9rem;border-bottom:1px solid var(--rule-light);font-family:var(--font-heading);font-weight:500;line-height:normal}.species-list__family-title>span{padding-bottom:1rem;border-bottom:5px solid var(--rule-light)}.species-list__family+.species-list__family{margin-top:3%}.species-list__subfamily-title{margin:0.6em 0 0.8em;font-family:var(--font-heading);font-weight:500}.species-list__items+.species-list__subfamily-title{margin-top:0.6em}.species-list__appendix{margin:1em 0 0.5em;padding:2% 0;font-family:var(--font-heading);font-size:250%;font-weight:300;line-height:1.2;color:var(--gray-300);background-image:linear-gradient(to right,var(--hr-dot) 1px,transparent 1px);background-size:3px 1px;background-repeat:repeat-x;background-position:0 100%}.species-list__appendix small{display:block;font-size:60%;line-height:1.2}.featured{position:relative;margin:0 0 3%;clear:both;--featured-height:clamp(18rem,28vw,35rem);--featured-gap:3px}.featured__track{display:flex;gap:var(--featured-gap);margin:0;padding:0;list-style:none;overflow-x:auto;scroll-snap-type:x 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#fff;background:var(--nav-panel-bg);color:#fff;opacity:0.8;cursor:pointer;display:none}.featured__btn svg{width:2.4rem;height:2.4rem}.featured__btn:focus-visible,.featured__btn:hover{opacity:0.55}.featured__btn--prev{left:0}.featured__btn--next{right:0;border-right:0;border-left:var(--featured-gap) solid #fff}.featured.has-overflow .featured__btn{display:inline-flex;align-items:center;justify-content:center}@media (prefers-reduced-motion:reduce){.featured__track{scroll-behavior:auto}.featured__caption{transition:none}}.rotator{padding:1rem 0}.rotator__panel{margin:0;background-color:transparent;border:0}.rotator.is-ready .rotator__panels{display:grid;margin-bottom:1rem}.rotator.is-ready .rotator__panel{grid-area:1/1;opacity:0;visibility:hidden;transition:opacity 0.6s ease,visibility 0s linear 0.6s}.rotator.is-ready .rotator__panel.is-active{opacity:1;visibility:visible;transition:opacity 0.6s ease,visibility 0s}@media (prefers-reduced-motion:reduce){.rotator.is-ready .rotator__panel{transition:none}}.rotator__actions{display:flex;flex-wrap:wrap;justify-content:flex-end;gap:var(--space-2);margin:0}.callout{margin-bottom:3rem;padding:4% 5%;background:var(--well-bg)}.callout>h2,.callout>h3{margin-top:0}.callout>:last-child{margin-bottom:0}.callout__media{float:left;margin:0.5rem 1rem 0 0}.callout__actions{text-align:right}.home-sidebar .widget-box{margin-bottom:var(--space-5)}.lightbox{--lb-duration:0.3s;width:fit-content;max-width:none;max-height:none;margin:auto;padding:0;border:0;border-radius:var(--radius);background:#fff;color:var(--color-text);box-shadow:0 1rem 4rem rgba(0,0,0,0.45);overflow:visible;opacity:0;transform:scale(0.96);transition:opacity var(--lb-duration) ease,transform var(--lb-duration) ease,overlay var(--lb-duration) ease allow-discrete,display var(--lb-duration) ease allow-discrete}.lightbox[open]{opacity:1;transform:none}@starting-style{.lightbox[open]{opacity:0;transform:scale(0.96)}}.lightbox::backdrop{background:rgba(0,0,0,0);transition:background var(--lb-duration) ease,overlay var(--lb-duration) ease allow-discrete,display var(--lb-duration) ease allow-discrete}.lightbox[open]::backdrop{background:rgba(0,0,0,0.8)}@starting-style{.lightbox[open]::backdrop{background:rgba(0,0,0,0)}}.lightbox__box{padding:1rem}.lightbox__stage{position:relative;width:40rem;height:30rem;max-width:94vw;background:var(--neutral-900);overflow:hidden;transition:width var(--lb-duration) ease,height var(--lb-duration) ease}.lightbox__img{position:absolute;inset:0;width:100%;height:100%;object-fit:contain;opacity:0;transition:opacity 0.35s ease}.lightbox__img.is-shown{opacity:1}.lightbox.is-loading .lightbox__stage:after{content:"";position:absolute;inset:0;background:rgba(0,0,0,0.25)}.lightbox__nav{position:absolute;top:50%;width:4.4rem;height:4.4rem;padding:0;border:0;border-radius:50%;background:rgba(0,0,0,0.45);color:#fff;font-size:3.2rem;line-height:1;cursor:pointer;transform:translateY(-50%);opacity:0;transition:opacity 0.25s ease,background 0.2s ease}.lightbox__nav span{position:relative;top:-0.2rem}.lightbox__nav--prev{left:1rem}.lightbox__nav--next{right:1rem}.lightbox__nav:focus-visible,.lightbox__nav:hover{background:rgba(0,0,0,0.75);opacity:1}.lightbox__nav:focus-visible,.lightbox__stage:hover .lightbox__nav{opacity:1}.lightbox--single .lightbox__nav{display:none}.lightbox__bar{display:flex;align-items:center;gap:var(--space-3);min-height:3.6rem;padding:0.8rem 0 0;font-size:1.4rem;color:var(--color-muted)}.lightbox__text{flex:1 1 auto;min-width:0}.lightbox__count{flex:none;white-space:nowrap}.lightbox__close{flex:none;width:2.8rem;height:2.8rem;padding:0;border:0;border-radius:50%;background:var(--neutral-200);color:var(--color-heading);font-size:2rem;line-height:1;cursor:pointer}.lightbox__close:focus-visible,.lightbox__close:hover{background:var(--color-primary);color:#fff}@media (max-width:600px){.lightbox__box{padding:0.6rem}.lightbox__nav{opacity:1;width:3.6rem;height:3.6rem;font-size:2.6rem}}@media (prefers-reduced-motion:reduce){.lightbox,.lightbox::backdrop,.lightbox__img,.lightbox__nav,.lightbox__stage{transition:none}}body:has(dialog.lightbox[open]){overflow:hidden}.book-price{margin:var(--space-3) 0;padding-bottom:var(--space-3);border-bottom:1px solid var(--rule-light);font-family:var(--font-heading);font-size:1.8rem;font-weight:500}.skip-link{position:absolute;top:-100%;left:var(--space-2);z-index:2000;padding:var(--space-2) var(--space-3);background:var(--color-primary);color:#fff;text-decoration:none;border-radius:0 0 var(--radius) var(--radius)}.skip-link:focus{top:0;outline:2px solid #fff;outline-offset:-4px}.site-header{position:relative;z-index:999;margin-bottom:0;background:var(--header-bg)}.site-header__top{background:#fff}.site-header__tools{display:flex;justify-content:flex-end;align-items:center;gap:var(--space-4);min-height:4.5rem}.site-header{}.site-header__bar{display:flex;align-items:center;justify-content:space-between;gap:var(--space-4)}@media print{.site-header{display:none}}.site-logo{display:block;flex:none;margin:0.5rem 0 1rem;line-height:0}.site-logo img{display:block;width:243px;height:81px;max-width:100%}.print-logo{display:none}.site-search{display:flex;align-items:center;gap:var(--space-2);margin:0}.site-search__label{margin:0;line-height:1;color:var(--search-trigger-text);font-size:85%;font-weight:700;text-transform:uppercase;letter-spacing:0.04em}.site-search .search-combo>input{width:18rem}.site-search .search-combo>.button,.site-search .search-combo>button,.site-search .search-combo>input{height:3rem}.search-toggle{display:none}.social-links{display:flex;align-items:center;gap:var(--space-3);margin:0;padding:0;list-style:none}.social-links a{display:inline-flex;align-items:center;justify-content:center;width:2.4rem;height:2.4rem;color:var(--search-trigger-text);text-decoration:none}.social-links a:focus-visible,.social-links a:hover{color:var(--color-link)}.social-links svg{display:block}.site-header__top .social-links{padding-right:var(--space-4);border-right:1px solid var(--rule-light);height:4.5rem}.nav-toggle{display:none}.site-nav__list,.site-nav__sub{margin:0;padding:0;list-style:none}.site-nav__list{display:flex;align-items:stretch}.site-nav__item{position:relative;margin-left:2px;background:var(--header-bg)}.site-nav{}.site-nav__link,.site-nav__toggle{display:flex;align-items:center;gap:0.8rem;height:9.5rem;padding:0 1.2rem;border:0;background:transparent;color:var(--nav-text);font:inherit;font-size:110%;text-transform:lowercase;text-decoration:none;cursor:pointer}@media (min-width:768px) and (max-width:899px){.site-nav__link,.site-nav__toggle{gap:0.5rem;padding:0 0.7rem;font-size:100%}}.site-nav__link:focus-visible,.site-nav__link:hover,.site-nav__link[aria-expanded=true],.site-nav__toggle:focus-visible,.site-nav__toggle:hover,.site-nav__toggle[aria-expanded=true]{color:#fff;background:var(--nav-hover)}.site-nav{}.site-nav__toggle:after{content:"";width:0.5rem;height:0.5rem;border-right:1px solid currentColor;border-bottom:1px solid currentColor;transform:translateY(-0.15rem) rotate(45deg);opacity:0.8}.site-nav__sub{display:none;position:absolute;top:100%;left:0;z-index:100;min-width:19.5rem;background:var(--nav-panel-bg)}.site-nav__sub li{border-bottom:1px solid var(--nav-sub-rule)}.site-nav__sub li:last-child{border-bottom:0}.site-nav__sub a{display:block;padding:0.7rem 1rem;background:var(--nav-sub-bg);color:#fff;line-height:1.5;text-transform:lowercase;text-decoration:none}.site-nav__sub a:focus-visible,.site-nav__sub a:hover{background:var(--nav-hover)}.site-nav{}.site-nav__item:hover>.site-nav__sub,.site-nav__toggle[aria-expanded=true]+.site-nav__sub{display:block}.site-nav{}.site-nav__item:nth-last-child(-n+2)>.site-nav__sub{left:auto;right:0}@media print{.site-nav{display:none}}@media (max-width:767px){.site-header__bar,.site-header__top .wrap{width:100%}.site-header__tools{box-sizing:border-box;justify-content:flex-start;gap:var(--space-2);min-height:5rem;padding:0.5rem}.site-header__top .social-links{display:none}.site-header__bar{flex-wrap:wrap;justify-content:center;gap:0}.site-logo{margin:0.8rem auto}.site-logo img{width:180px;height:60px}.nav-toggle{display:inline-flex;align-items:center;justify-content:center;flex:none;width:4rem;height:4rem;padding:0;border:0;background:var(--nav-hover);color:#fff;cursor:pointer}.nav-toggle__bars{position:relative;display:block;width:2rem;height:2px;background:currentColor;transition:background 120ms}.nav-toggle__bars:after,.nav-toggle__bars:before{content:"";position:absolute;left:0;width:100%;height:2px;background:currentColor;transition:transform 120ms}.nav-toggle__bars:before{top:-6px}.nav-toggle__bars:after{top:6px}.nav-toggle[aria-expanded=true] .nav-toggle__bars{background:transparent}.nav-toggle[aria-expanded=true] .nav-toggle__bars:before{transform:translateY(6px) rotate(45deg)}.nav-toggle[aria-expanded=true] .nav-toggle__bars:after{transform:translateY(-6px) rotate(-45deg)}.search-toggle{display:inline-flex;align-items:center;justify-content:center;flex:none;width:4rem;height:4rem;padding:0;border:0;background:transparent;color:var(--search-trigger-text);cursor:pointer}.search-toggle[aria-expanded=true]{color:var(--color-link)}.site-search{flex:1 1 auto}.site-search:not(.is-open){display:none}.site-search__label{display:none}.site-search .search-combo{flex:1 1 auto}.site-search .search-combo>input{width:auto}.site-nav{flex-basis:100%;background:var(--header-bg);border-top:1px solid rgba(163,176,187,0.4)}.site-nav:not(.is-open){display:none}.site-nav__list{flex-direction:column;padding:1rem 0}.site-nav__item{margin:0}.site-nav__link,.site-nav__toggle{width:100%;height:auto;padding:1rem 2rem 0.5rem;font-size:100%;text-transform:uppercase;justify-content:space-between}.site-nav__toggle[aria-expanded=true]:after{transform:translateY(0.15rem) rotate(-135deg)}.site-nav__sub{position:static;min-width:0;background:transparent}.site-nav__sub li{border:0}.site-nav__sub a{padding:0.8rem 0 0.5rem 3rem;background:var(--header-bg);text-transform:none}.site-nav{}.site-nav__item:hover>.site-nav__sub{display:none}.site-nav__toggle[aria-expanded=true]+.site-nav__sub{display:block}}.site-footer{clear:both;padding-top:2%;border-top:3px solid var(--nav-panel-bg);background:var(--header-bg);color:var(--nav-text)}.site-footer a{color:var(--footer-link);text-decoration:none}.site-footer a:focus-visible,.site-footer a:hover{text-decoration:underline}.site-footer p,.site-footer ul{color:var(--nav-text)}.site-footer__grid{margin-bottom:2%}.site-footer__section{padding-bottom:1.5rem}.site-footer__heading{margin:0 0 1rem;padding:0 0 0.5rem;border-bottom:1px solid var(--nav-panel-bg);font-family:var(--font-heading);font-size:120%;font-weight:500;color:var(--nav-text)}.site-footer .social-links{margin-bottom:var(--space-2)}.site-footer .social-links a{color:var(--footer-link)}.site-footer .social-links a:focus-visible,.site-footer .social-links a:hover{color:#fff}.site-footer{}.site-footer .footer-search{margin:0 0 1.5rem}.site-footer .footer-search__label{position:absolute;width:1px;height:1px;margin:-1px;overflow:hidden;clip:rect(0 0 0 0);white-space:nowrap}.site-footer .footer-search .search-combo{max-width:32rem}.site-footer{}.site-footer__thumbs{display:flex;flex-wrap:wrap;gap:1rem;margin:0;padding:0;list-style:none}.site-footer__thumbs li{width:50px;height:50px}.site-footer__thumbs img{display:block;width:50px;height:50px;border:1px solid var(--gray-500)}.site-footer__thumbs a:focus-visible img,.site-footer__thumbs a:hover img{border-color:var(--gray-300)}.site-footer{}.site-footer__books{margin:0;padding:0;list-style:none}.site-footer__books li{padding:0.5rem 0}.site-footer__books li:first-child{padding-top:0}.site-footer__books .meta{display:block;font-size:70%;text-transform:uppercase;color:var(--footer-meta)}.site-footer__credits{padding:1rem;background:var(--footer-credits-bg);font-size:80%}.site-footer__credits ul{display:flex;flex-wrap:wrap;justify-content:flex-end;margin:0;padding:1rem 0;list-style:none}.site-footer__credits li{padding:0 1rem;border-right:1px solid var(--footer-credits-rule);color:var(--footer-text);text-transform:uppercase}.site-footer__credits li:last-child{border-right:0;padding-right:0}.site-footer__credits a{color:var(--footer-text);text-transform:uppercase}.site-footer__credits a:focus-visible,.site-footer__credits a:hover{color:#fff}@media print{.site-footer{display:none}}.back-to-top{position:fixed;right:1rem;bottom:1rem;z-index:1000;display:flex;align-items:center;justify-content:center;width:5rem;height:5rem;background:var(--nav-hover);color:#fff;text-decoration:none;opacity:0;visibility:hidden;transition:opacity 200ms,visibility 200ms}.back-to-top.is-visible{opacity:1;visibility:visible}.back-to-top:focus-visible,.back-to-top:hover{background:var(--footer-link);color:#fff}.back-to-top svg{display:block}@media print{.back-to-top{display:none}}@media (prefers-reduced-motion:no-preference){html{scroll-behavior:smooth}}@media print{.print-logo{display:block;margin:0 auto}}.well{min-height:20px;margin-bottom:20px;padding:2% 4%;background:var(--well-bg);border:0;border-radius:0;box-shadow:none}.well>:last-child{margin-bottom:0}.dropcap{display:block;float:left;margin:2px 5px 0 0;padding:0;font-size:45px;font-style:normal;line-height:45px;color:var(--color-heading);text-transform:uppercase}@media (max-width:500px){.dropcap{font-size:30px;line-height:30px}}img.align_left{float:left;margin:1% 3% 1% 0}img.align_right{float:right;margin:1% 0 1% 3%}img.align_center{margin-inline:auto}.btn{display:inline-flex;align-items:center;gap:var(--space-1);padding:var(--space-1) var(--space-3);border:1px solid var(--color-primary);border-radius:var(--radius);background:var(--color-primary);color:#fff;font:inherit;line-height:1.5;text-decoration:none;text-shadow:none;box-shadow:none;cursor:pointer}.btn:focus-visible,.btn:hover{background:var(--color-link);border-color:var(--color-link);color:#fff;text-decoration:none}.btn.btn-very-subtle{background:var(--neutral-200);border-color:var(--neutral-400);color:var(--color-text)}.btn.btn-very-subtle:focus-visible,.btn.btn-very-subtle:hover{background:var(--color-muted);border-color:var(--color-muted);color:#fff}.btn svg{flex:none}.Inputfield{margin-bottom:20px}.InputfieldContent input,.InputfieldContent textarea{width:98%}.InputfieldContent p.help-block{margin-top:-14px;margin-bottom:20px;padding-left:5px;font-size:120%;background-color:var(--info-bg)}.help-block{display:block;margin-bottom:10px;color:var(--color-muted-text)}p.text-danger{margin-bottom:0;color:var(--danger)}.FormBuilderErrors{margin-bottom:10px;padding:10px;color:var(--danger);background-color:var(--danger-bg);border:1px solid var(--danger-border)}#wrap_Inputfield-{display:none}.distmap{box-sizing:border-box;width:100%;min-width:200px;min-height:400px;aspect-ratio:4/5;background:var(--gray-100);border:1px solid var(--distmap-border)}@supports not (aspect-ratio:1){.distmap{height:450px}}.distmap__caption{display:flex;flex-wrap:wrap;align-items:center;gap:0.5rem 1.2rem;padding:0.85rem 0.4rem 0;font-size:1.15rem;line-height:1.4}.distmap__total{font-weight:600}.distmap__legend{display:inline-flex;align-items:center;gap:0.4rem}.distmap__legend-label{margin-right:0.6rem}.distmap__swatch{display:inline-block;width:1.1rem;height:1.1rem;margin-right:0.35rem;border:1px solid rgba(0,0,0,0.25);vertical-align:-3px}.distmap__swatch--pre2000{background:var(--distmap-pre2000)}.distmap__swatch--post2000{background:var(--distmap-post2000)}.distmap__truncated{color:var(--warning-text);font-style:italic}.distmap__nbnlink{margin-left:auto}.distmap__empty{box-sizing:border-box;display:flex;flex-direction:column;align-items:center;justify-content:center;width:100%;min-height:400px;aspect-ratio:4/5;padding:2rem;background:var(--gray-100);border:1px solid var(--distmap-border);color:var(--distmap-empty-text);text-align:center}.distmap__empty p{max-width:32ch;margin:0.5em 0;line-height:1.4}.distmap__empty-icon{display:block;margin-bottom:0.75rem;color:var(--distmap-empty-icon)}.distmap__empty-icon svg{display:block}.distmap__empty-icon--spin svg{animation:distmap-spin 1s linear infinite}@keyframes distmap-spin{to{transform:rotate(360deg)}}@media (prefers-reduced-motion:reduce){.distmap__empty-icon--spin svg{animation:none}}.distmap__error{padding:1rem;color:var(--distmap-error);text-align:center}.leaflet-tooltip{font-size:0.85rem}.flightperiod{margin:1rem 0 1.25rem;max-width:360px}.flightperiod__divider{margin:0 0 10px}.flightperiod__title{margin:0 0 0.4rem;font-size:1.2rem;font-weight:600;color:var(--color-heading)}.flightperiod__chart{margin:0}.flightperiod__grid{display:grid;grid-template-columns:repeat(12,1fr);gap:1px;height:1.8rem;border:1px solid var(--neutral-500);background:#fff}.flightperiod__cell{--intensity:0;background:hsl(188,calc(40% + 35% * var(--intensity)),calc(94% - 50% * var(--intensity)));transition:background-color 200ms ease}.flightperiod__cell[data-zero]{background:var(--neutral-150)}.flightperiod__axis{display:grid;grid-template-columns:repeat(12,1fr);gap:0;margin-top:0.2rem;font-size:1rem;color:var(--neutral-600);text-align:center}.flightperiod__caption{margin-top:0.4rem;font-size:1.2rem;line-height:1.4;color:var(--neutral-900)}.flightperiod__caption a{color:inherit;text-decoration:underline}.flightperiod--limited .flightperiod__grid{opacity:0.4;filter:grayscale(1)}.flightperiod[data-palette=yellow-red] .flightperiod__cell{background:hsl(calc(60deg - 40deg * var(--intensity)),calc(80% + 10% * var(--intensity)),calc(85% - 35% * var(--intensity)))}.flightperiod[data-palette=monochrome] .flightperiod__cell{background:hsl(0,0%,calc(95% - 60% * var(--intensity)))}@media (max-width:320px){.flightperiod__axis span:not(:nth-child(3n+1)){visibility:hidden}}.flightperiod__hidden{position:absolute!important;width:1px;height:1px;padding:0;margin:-1px;overflow:hidden;clip:rect(0 0 0 0);white-space:nowrap;border:0}@media (prefers-reduced-motion:reduce){.flightperiod__cell{transition:none}}@media (prefers-contrast:more){.flightperiod__cell{background:#fff;background-image:repeating-linear-gradient(45deg,#000 0,#000 calc(1px + 4px * var(--intensity)),transparent calc(1px + 4px * var(--intensity)),transparent 6px)}}@media print{.flightperiod__cell{background:hsl(0,0%,calc(95% - 60% * var(--intensity)))!important;background-image:none!important}}.idwiz-intro{max-width:46rem;margin:0.5rem 0 0;font-size:17px}#idwiz{position:relative;margin:1.5rem 0 3rem;font-size:16px;line-height:1.45}.idwiz-loading{display:none;position:absolute;inset:0;z-index:20;background:rgba(255,255,255,0.66)}#idwiz.is-loading .idwiz-loading{display:block}.idwiz-spinner{position:fixed;top:45%;left:50%;width:44px;height:44px;margin:-22px 0 0 -22px;border:4px solid rgba(54,128,147,0.25);border-top-color:var(--color-primary);border-radius:50%;animation:idwiz-spin 0.7s linear infinite}@keyframes idwiz-spin{to{transform:rotate(360deg)}}.idwiz-grid{display:flex;flex-wrap:wrap;gap:2rem;align-items:flex-start}.idwiz-panel{flex:1 1 360px;min-width:0}.idwiz-results{flex:1 1 320px;min-width:0;position:sticky;top:1rem}@media (max-width:760px){.idwiz-results{position:static}}.idwiz-progress{text-transform:uppercase;letter-spacing:0.06em;font-size:13px;color:var(--color-muted-text);margin:0 0 0.25rem}.idwiz-q{margin:0 0 0.25rem;font-size:26px}.idwiz-req{font-size:14px;color:var(--color-muted-text);font-weight:400}.idwiz-results-head,.idwiz-step-head{min-height:76px}.idwiz-step-head{display:flex;justify-content:space-between;align-items:flex-start;gap:0.5rem 1.5rem;flex-wrap:wrap}.idwiz-step-head .idwiz-nav{flex:0 0 auto;justify-content:flex-end}.idwiz-step-head-text{flex:1 1 240px;min-width:0}.idwiz-hint,.idwiz-notice{margin:0 0 0.75rem;color:var(--color-muted-text)}.idwiz-notice{color:var(--danger-strong);font-weight:600}.idwiz-cards{display:grid;grid-template-columns:repeat(auto-fill,minmax(140px,1fr));gap:0.75rem;margin:0 0 1.25rem}.idwiz-card{position:relative;display:flex;flex-direction:column;border:1px solid var(--neutral-400);border-radius:3px;padding:0.5rem;cursor:pointer;background:#fff;transition:border-color 0.12s ease,box-shadow 0.12s ease}.idwiz-card:hover{border-color:var(--color-link)}.idwiz-card{}.idwiz-card input{position:absolute;top:8px;left:8px;width:22px;height:22px;margin:0;z-index:2;cursor:pointer;appearance:auto}.idwiz-card input[type=checkbox]{-webkit-appearance:checkbox}.idwiz-card.is-checked{border-color:var(--color-primary);box-shadow:0 0 0 2px var(--color-primary) inset}.idwiz-card:focus-within{outline:2px solid var(--color-link);outline-offset:2px}.idwiz-card-img{width:100%;height:104px;margin-top:6px;object-fit:cover;border-radius:2px;display:block;margin-bottom:0.45rem;background:var(--neutral-200)}.idwiz-card-label{font-weight:700;font-size:17px;line-height:1.25}.idwiz-card-help{font-size:15px;color:var(--color-muted-text);margin-top:0.25rem;line-height:1.35}.idwiz-nav{display:flex;flex-wrap:wrap;gap:0.5rem;align-items:center}.idwiz-btn-link{background:transparent;border:0;color:var(--color-link);font-size:16px;cursor:pointer;padding:12px 6px}.idwiz-btn-link:hover{text-decoration:underline}.idwiz-results-head{display:flex;justify-content:space-between;align-items:flex-start;gap:0.5rem 1rem;border-bottom:2px solid var(--neutral-400);padding-bottom:0.4rem;margin-bottom:0.75rem}.idwiz-results-head-text{min-width:0}.idwiz-results-restart{flex:0 0 auto;margin:0}.idwiz-count-line{margin:0}.idwiz-count{font-size:30px;font-weight:700;color:var(--color-primary)}.idwiz-count-label{font-size:16px;color:var(--color-muted-text)}.idwiz-count-sub{margin:0.15rem 0 0;font-size:13px;color:var(--color-muted-text)}.idwiz-nomatch,.idwiz-relaxed{color:var(--danger-strong);font-size:0.85rem;margin:0 0 0.75rem}.idwiz-nomatch{margin-top:0.5rem}.idwiz-tiles{list-style:none;margin:0;padding:0;display:grid;grid-template-columns:repeat(auto-fill,minmax(120px,1fr));gap:0.6rem}.idwiz-tile{transition:opacity 0.2s ease}.idwiz-tile a{display:block;text-decoration:none;color:inherit}.idwiz-tile .idwiz-tile-img,.idwiz-tile img{display:block;width:100%;height:auto;border-radius:2px}.idwiz-tile.is-partial{opacity:0.5}.idwiz-tile.is-faded{opacity:0.28}.idwiz-tile-noimg{height:90px;background:var(--neutral-200) repeating-linear-gradient(45deg,var(--neutral-200),var(--neutral-200) 8px,var(--neutral-250) 8px,var(--neutral-250) 16px)}.idwiz-tile-name{display:block;font-size:15px;line-height:1.25;margin-top:0.3rem}.idwiz-empty{color:var(--color-muted-text)}.idwiz-placeholder{color:var(--color-muted-text);border:1px dashed var(--neutral-400);border-radius:3px;padding:1.25rem 1rem;margin:0;font-size:15px;line-height:1.5}.idwiz-done{max-width:40rem}.crumbs{margin-bottom:3%;background:var(--gray-200);font-family:var(--font-heading)}.crumbs__inner{display:flex;flex-wrap:wrap;justify-content:space-between;align-items:baseline;gap:0 var(--space-4);padding:1rem 0}.crumbs p{margin:0}.crumbs__taxa{font-size:1.95rem}.crumbs__nav{font-size:1.65rem;margin-left:auto}.crumbs__sep{color:var(--color-heading)}.crumbs a{color:var(--color-breadcrumb-link)}.crumbs a:hover{color:var(--color-link)}@media (max-width:767px){.crumbs__inner{flex-direction:column;align-items:stretch;gap:0.4rem}.crumbs__taxa{font-size:2.08rem}.crumbs__nav{font-size:1.76rem;text-align:right;margin-left:0}}.species-layout{display:grid;grid-template-columns:minmax(0,1fr);gap:var(--gutter);align-items:start}@media (min-width:768px){.species-layout{grid-template-columns:5fr 7fr;gap:2.56%}}.species-media{container-type:inline-size;min-width:0}.species-text{min-width:0;padding-bottom:2%}.species-header{display:flex;flex-direction:column;margin-bottom:1rem;padding-bottom:1rem;background-image:linear-gradient(to right,var(--hr-dot) 1px,transparent 1px);background-size:3px 1px;background-repeat:repeat-x;background-position:0 100%}.species-header>*{margin:0}.species-checklist{order:-1;margin-top:1.2rem;font-family:var(--font-heading);font-size:3rem;font-weight:400;line-height:1.2;color:var(--color-heading)}.species-checklist small{font-size:60%}@media (max-width:767px){.species-checklist{font-size:2.1rem;font-weight:500}}.species-title{margin-bottom:1.2rem;font-family:var(--font-heading);font-size:3rem;font-weight:400;line-height:1.2;color:var(--color-heading)}.species-title strong{font-weight:600}.species-title em{font-weight:300}@media (max-width:767px){.species-title{font-size:3.2rem}}.species-title--micro{line-height:1.1}.species-title--micro strong{font-size:2.3rem;font-weight:600}.species-author{font-family:var(--font-heading);font-size:2.4rem;font-weight:400;line-height:1.2;color:var(--color-heading)}@media (max-width:767px){.species-author{font-size:1.9rem;font-weight:500}}.species-synonyms{font-size:1.4rem;margin-top:0.4rem}.species-description>:first-child{margin-top:0}.references p{margin-bottom:0}#imgcontainer{margin:2% 0 0}#imgcontainer #imgcontainerhead{margin:0}#imgcontainer #tab{line-height:1.2;padding-bottom:0.4rem}#imgcontainer img{display:block;max-width:100%;height:auto;margin-inline:auto}#imgcontainer .species-figure__link{display:block;cursor:zoom-in}#imgcontainer figcaption{position:relative;display:flex;justify-content:space-between;gap:var(--space-2);padding:1rem;background:var(--gray-50);font-size:90%}#imgcontainer figcaption .contribphotos{flex:none}#imgcontainer .map{min-height:26rem;margin-top:1rem;padding:2rem;background-color:rgba(199,207,214,0.4)}.species-more{margin:1rem 0 0;border-bottom:0}.species-more>summary{text-align:right}.species-more__body{padding-bottom:var(--space-3)}.species-more iframe{width:100%}.species-related{display:flex;flex-wrap:wrap;justify-content:space-between;gap:0 var(--gutter)}.related{clear:both;flex:1 1 30rem;margin-top:1rem;padding-top:1.5rem;background-image:linear-gradient(to right,var(--hr-dot) 1px,transparent 1px);background-size:3px 1px;background-repeat:repeat-x}.related__list{display:flex;flex-wrap:wrap;gap:1rem;margin:0 0 1rem;padding:0;list-style:none}.related__item{flex:none;width:7.5rem;height:7.5rem}.related__item a{display:block;width:100%;height:100%;overflow:hidden;border:1px solid #fff}.related__item a:focus-visible,.related__item a:hover{border-color:var(--gray-600)}.related__item img{display:block;width:7.5rem;height:7.5rem}.related .section-title{margin-bottom:1.5rem}.references p{margin-bottom:0.5rem}@media print{.crumbs,.species-more,.species-related{display:none}}}@layer utilities{.visually-hidden:not(:focus):not(:active){position:absolute;width:1px;height:1px;margin:-1px;padding:0;overflow:hidden;clip:rect(0 0 0 0);clip-path:inset(50%);white-space:nowrap;border:0}.text-right{text-align:right}.clear{clear:both}svg.flip-h{transform:scaleX(-1)}}
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